Alternate Spatial Lotka-Volterra Equations
• Let Si(x,y) be density of the ith species (rabbits, trees, seeds, …)
• dSi / dt = riSi(1 - Si - ΣaijSj)
2-D grid:
  S = Sx-1,y + Sx,y-1
+ Sx+1,y + Sx,y+1 + aSx,y 
j¹i
where